string: It has also been shown that boule functions in spermatogenesis by controlling the translation of the meiotic CDC25 cell cycle regulators, especially CDC25A (Xu et al., 2001; VanGompel and Xu, 2010), which is expressed in the testes and might play a role in the mitotic or meiotic regulation via G1/S transition and M-phase exit (Jinno et al.