string: PCR amplification and DNA sequencing Eight loci including the partial rDNA-ITS (ITS), ac t in (ACT ) , ch i t in syn thase 1 (CHS-1 ) , glyceraldehyde-3-phosphate dehydrogenase (GAPDH), β-tubulin (TUB2), Histone (CYLH3), glutamine synthetase (GS) and ApMat were amplified and sequenced using the primer pairs ITS-1 F (Gardes and Bruns 1993) /ITS-4 (White et al.