text: …studies where have shown that the heterologous expression
  1 3
  of R2R3 and R3-MYB transcription factors is capable to suppress flavonoids in model plants such as tobacco and Arabidopsis (Aharoni et al. 2001; Matsui et al. 2008; Zhu et al. 2009, 2013; Albert et al. 2011; Nakatsuka et al. 2013).